What Animal Has As Many Neurons In Brain As Humans Per Weight

Brief comparisons of number of whole brain neurons (summit) and cerebral cortex neurons (bottom) amongst vi mammals.

The following are two lists of animals ordered by the size of their nervous organisation. The start list shows number of neurons in their unabridged nervous system, indicating their overall neural complexity. The 2d list shows the number of neurons in the structure that has been establish to be representative of animal intelligence.^[one] The human brain contains 86 billion neurons, with 16 billion neurons in the cerebral cortex.^{[2] [1]}

Overview [edit]

Neurons are the cells that transmit information in an animal's nervous system then that it can sense stimuli from its environment and behave accordingly. Not all animals have neurons; Trichoplax and sponges lack nerve cells altogether.

Neurons may be packed to form structures such as the brain of vertebrates or the neural ganglions of insects.

The number of neurons and their relative abundance in different parts of the brain is a determinant of neural office and, consequently, of beliefs.

Whole nervous organization [edit]

All numbers for neurons (except Caenorhabditis and Ciona), and all numbers for synapses (except Ciona) are estimations.

Name	Neurons in the brain & whole nervous system	Synapses	Details	Image	Source
Sponge	0				[3]
Trichoplax	0		Despite no nervous system, it exhibits coordinated feeding and response behaviors.[iv]		[5]
Asplanchna brightwellii (rotifer)	nearly 200		Brain only		[half-dozen]
Tardigrade	most 200		Brain only		[7]
Ciona intestinalis larva (sea eject)	231	8,617 (key nervous organization just)			[8] [nine]
Caenorhabditis elegans (roundworm)	302	~7,500	is the but organism to take its whole connectome (neuronal "wiring diagram") completed.[10] [11] [12]		[13]
Jellyfish	v,600		Hydra vulgaris (H. attenuate)		[14]
Megaphragma mymaripenne	7,400				[15]
Box jellyfish	8,700–17,500		adult Tripedalia cystophora (eight mm bore) – does not include 1000 neurons in each of the four rhopalia		[16]
Medicinal leech	10,000				[17]
Pond snail	xi,000				[18]
Sea slug	18,000				[xix]
Amphioxus	20,000		central nervous system only		[20] [21]
Larval zebrafish	100,000				[22]
Lobster	100,000				[23]
Fruit fly	100,000	< x,000,000 [ citation needed ]			[24]
Wandering spider	100,000				[25]
Calliopsis (bee)	234,000				[26]
Pismire	250,000		Varies per species		[27]
Perdita (genus)	275,000				[26]
Melissodes	495,000				[26]
Bombus impatiens	557,000				[26]
Western honey bee	613,000				[26]
Honey bee	960,000	~i×10 ^ 9			[28]
Cockroach	1,000,000				[29]
Coconut crab	>1,000,000		c. 1,000,000 interneurons are dedicated to analysing olfactory input lonely.		[30]
California carpenter bee	1,180,000				[26]
Steudner's dwarf gecko	1,771,000				[31]
Brown anole	two,792,000				[31]
Mochlus sundevallii	iii,049,000				[31]
Peloponnese slowworm	three,713,000				[31]
Mutual business firm gecko	iii,988,000				[31]
Takydromus sexlineatus	4,021,000				[31]
Anolis cristatellus	iv,270,000				[32]
Papua snake cadger	four,271,000				[31]
Guppy	4,300,000				[33]
Natal Midlands dwarf chameleon	4,305,000				[31]
Acontias percivali	4,340,000				[31]
Sand lizard	4,341,000				[31]
Ocelot gecko	4,420,000				[34]
Darevskia raddei	iv,765,000				[31]
Anolis evermanni	iv,920,000				[32]
Echis carinatus	4,951,000				[31]
Cerastes cerastes	4,996,000				[31]
Tenerife gecko	v,001,000				[31]
Draco sumatranus	v,174,000				[31]
Blue-throated keeled lizard	5,269,000				[31]
Crested gecko	5,417,000				[31]
Lacerta strigata	five,529,000				[31]
San Francisco garter snake	5,663,000				[31]
Scarlet-eyed crocodile skink	5,697,000				[31]
Emoia cyanura	v,733,000				[31]
East African spiny-tailed lizard	five,756,000				[31]
Chalcides ocellatus	five,774,000				[31]
Cylindrophis ruffus	5,779,000				[31]
True cat gecko	5,964,000				[31]
Aspidoscelis deppii	5,968,000				[31]
Brown water ophidian	5,995,000				[31]
Aspidelaps lubricus	6,020,000				[31]
Scincus scincus	6,284,000				[31]
Montivipera xanthina	6,677,000				[31]
Hispaniolan curlytail lizard	seven,063,000				[31]
Sceloporus malachiticus	7,149,000				[31]
Crotalus durissus	7,263,000				[31]
Agama agama	7,455,000				[31]
Agama aculeata	vii,631,000				[31]
Gekko kuhli	7,659,000				[31]
Leopard gecko	8,081,000				[31]
Latastia longicaudata	8,099,000				[31]
Razor-backed musk turtle	8,389,000				[31]
Greek tortoise	8,520,000				[31]
Phelsuma grandis	8,623,000				[31]
Acanthocercus atricollis	8,650,000				[31]
Tokay gecko	eight,892,000				[31]
Russian tortoise	nine,008,000				[31]
Marginated tortoise	9,074,000				[31]
Psammophis elegans	9,170,000				[31]
Xenopeltis unicolor	9,293,000				[31]
Zonosaurus Karsteni	ix,538,000				[31]
Oplurus quadrimaculatus	9,565,000				[31]
Malagasy giant chameleon	9,751,000				[31]
Ahaetulla prasina	nine,767,000				[31]
Dasia olivacea	ix,785,000				[31]
Adult zebrafish	~10,000,000		cells (neurons + other)		[35]
Timon tangitanus	10,619,000				[31]
Corn snake	10,629,000				[31]
Acanthosaura capra	10,724,000				[31]
Gallotia galloti	10,903,000				[31]
Eutropis multifasciata	10,944,000				[31]
Rainbow boa	11,083,000				[31]
East African black mud turtle	11,285,000				[31]
Indochinese spitting cobra	eleven,306,000				[31]
Cyrtodactylus irianjayaensis	11,415,000				[31]
Uromastyx ornata	eleven,438,000				[31]
Yellow-throated plated cadger	11,847,000				[31]
Meller's chameleon	12,035,000				[31]
Twist-necked turtle	12,171,000				[31]
Western whiptail	12,269,000				[31]
Gehyra vorax	12,388,000				[31]
White-lipped mud turtle	12,671,000				[31]
Painted wood turtle	12,677,000				[31]
Sheltopusik	12,830,000				[31]
Standing'south day gecko	thirteen,223,000				[31]
Common snapping turtle	13,681,000				[31]
Painted turtle	14,302,000				[31]
Ocellated cadger	14,593,000				[31]
Oplurus cuvieri	14,627,000				[31]
African spurred tortoise	14,986,000				[31]
Boa imperator	15,194,000				[31]
Red-eared slider	15,388,000				[31]
Plumed basilisk	fifteen,536,000				[31]
Ruby-bellied short-necked turtle	15,787,000				[31]
Frog	xvi,000,000				[36]
Corallus hortulana	sixteen,303,000				[31]
Blueish-tongued skink	xvi,376,000				[31]
Gran Canaria behemothic lizard	xviii,038,000				[31]
Chinese h2o dragon	eighteen,301,000				[31]
Chlamydosaurus	18,641,000				[31]
Northern ophidian-necked turtle	19,040,000				[31]
Spiny-tailed monitor	nineteen,771,000				[31]
Boa constrictor	xix,781,000				[31]
Ameiva ameiva	xix,835,000				[31]
Chinese softshell turtle	xx,697,000				[31]
Amboina box turtle	21,440,000				[31]
Alligator snapping turtle	22,230,000				[31]
Yellow anaconda	23,035,000				[31]
Mexican beaded lizard	23,681,000				[31]
Major skink	24,155,000				[31]
Sudan plated lizard	24,434,000				[31]
Mata mata	26,105,000				[31]
Naked mole-rat	26,880,000				[37]
Weber's sailfin cadger	27,245,000				[31]
Green iguana	29,098,000				[31]
Argentine black and white tegu	29,251,000				[31]
Cyclura nubila	thirty,820,000				[31]
Gold tegu	31,335,000				[31]
Varanus macraei	33,091,000				[31]
Fiddling gratuitous-tailed bat	35,000,000				[38]
Smoky shrew	36,000,000				[39]
Sand goanna	50,253,000				[31]
Brusk-tailed shrew	52,000,000				[39]
Squealer-nosed turtle	53,027,000				[31]
Common bent-wing bat	56,000,000				[38]
African sheath-tailed bat	59,000,000				[38]
Hottentot golden mole	65,000,000				[forty]
Woermann'due south bat	66,000,000				[38]
Zenker's fruit bat	66,000,000				[38]
Commerson's roundleaf bat	67,000,000				[38]
Rufous trident bat	lxx,000,000				[38]
House mouse	71,000,000	~1×x ^ 12			[41]
Asian water monitor	78,231,000				[31]
Male monarch quail	eighty,478,000				[31]
Nile crocodile	80,500,000				[42]
Heart-nosed bat	81,000,000				[38]
Diamond pigeon	87,879,000				[31]
Golden hamster	90,000,000				[41]
Short-palated fruit bat	99,000,000				[38]
Ansell's mole-rat	103,000,000				[43]
Mashona mole-rat	113,000,000				[43]
Common quail	117,760,000				[31]
Hairy-tailed mole	124,000,000				[twoscore]
Eastern rock elephant shrew	129,000,000				[xl]
Star-nosed mole	131,000,000				[39]
Zebra finch	131,000,000		Brain only		[44]
African collared dove	139,271,000				[31]
Silvery mole-rat	148,000,000				[43]
Northern bobwhite	148,399,000				[31]
Four-toed elephant shrew	157,000,000				[40]
Eurasian blackcap	157,000,000				[44]
Wahlberg'southward epauletted fruit bat	162,000,000				[38]
Goldcrest	164,000,000				[44]
Cape mole-rat	170,000,000				[43]
Grey partridge	170,287,000				[31]
Egyptian fruit bat	172,000,000				[38]
Mechow'due south mole-rat	174,000,000				[43]
Damaraland mole-rat	178,000,000				[43]
Franquet'southward epauletted fruit bat	186,000,000				[38]
Chocolate-brown rat	200,000,000	~4.48×10 ^ 11			[45]
Guyenne spiny rat	202,000,000				[40]
Eastern mole	204,000,000				[39]
Elegant crested tinamou	217,768,000				[31]
Cerise junglefowl	221,000,000				[44]
Neat tit	226,000,000				[44]
Green-rumped parrotlet	227,000,000				[44]
Quebracho crested tinamou	235,671,000				[31]
Guinea squealer	240,000,000				[41]
Chilean tinamou	241,374,000				[31]
Gray mouse lemur	254,710,000				[46]
Common wood pigeon	258,681,000				[31]
Common treeshrew	261,000,000				[47]
Hammer-headed bat	275,000,000				[38]
Wood duck	305,816,000				[31]
Dove	310,000,000		Encephalon only		[44]
Budgerigar	322,000,000				[44]
Pygmy falcon	350,367,000				[31]
Greatcoat dune mole-rat	361,000,000				[43]
Mallard	367,000,000				[31]
Eurasian teal	375,409,000				[31]
Common blackbird	379,000,000				[44]
Marbled duck	386,920,000				[31]
Reeves's pheasant	387,173,000				[31]
Eurasian sparrowhawk	401,751,000				[31]
Ferret	404,000,000				[48]
Cockatiel	453,000,000				[44]
Gray squirrel	453,660,000				[37]
Banded mongoose	454,000,000				[48]
Prairie canis familiaris	473,940,000				[37]
Cherry-red-breasted goose	482,705,000				[31]
Common starling	483,000,000				[44]
Wild turkey	492,873,000				[31]
European rabbit	494,200,000				[37]
Octopus	500,000,000				[49]
Western tree hyrax	505,000,000				[forty]
Eurasian pygmy owl	533,263,000				[31]
Indian peafowl	570,934,000				[31]
Victoria crowned dove	578,697,000				[31]
Little owl	585,923,000				[31]
Common kestrel	624,582,000				[31]
Common marmoset	636,000,000				[47]
Eastern rosella	642,000,000				[44]
Yellow-knobbed curassow	652,989,000				[31]
Barn owl	690,000,000				[44]
Monk parakeet	697,000,000				[44]
Greylag goose	738,232,000				[31]
Azure-winged magpie	741,000,000				[44]
Rock hyrax	756,000,000				[40]
Cat	760,000,000	~1×10 ^ xiii			[l]
Blackness-rumped agouti	857,000,000				[41]
Magpie	897,000,000				[44]
Common hill myna	906,000,000				[44]
Northern greater galago	936,000,000				[47]
Western jackdaw	968,000,000				[44]
Long-eared owl	991,310,000				[31]
Mute swan	1.001 ×109				[31]
Common buzzard	i.001 × 109				[31]
Black vulture	1.009 ×x9				[31]
Greater rhea	i.03× xnine				[31]
Eurasian jay	1.085×ten ^ 9				[44]
Alexandrine parakeet	1.096×x ^ 9				[44]
Raccoon dog	one.xvi×x ^ 9				[51]
Tanimbar corella	1.161×ten ^ 9				[44]
Saker falcon	1.204 ×109				[31]
Emu	1.335×x ^ nine				[44]
3-striped night monkey	1.468×x ^ 9				[47]
Rook	1.509×10 ^ 9				[44]
Gray parrot	1.566×10 ^ ix				[44]
Tawny owl	1.58×10ix				[31]
Capybara	1.6×10 ^ 9				[41]
Common ostrich	1.62× 10nine				[31]
White-tailed eagle	1.65 ×x9				[31]
Jackal	1.73×10 ^ 9				[51]
Snowy owl	ane.807×ten9				[31]
Eurasian hawkeye-owl	ane.897×10nine				[31]
Fox	2.11×10 ^ ix				[51]
Sulphur-crested cockatoo	2.122×x ^ nine				[44]
Raccoon	2.148×10 ^ nine				[48]
Kea	ii.149×ten ^ 9				[44]
Raven	2.171×x ^ 9		Brain simply		[44]
Domestic pig	2.22×10 ^ ix				[52]
Dog	two.253×10 ^ ix				[48]
Chihuahua (dog)	2.51×ten ^ 9				[51]
Dog (Chow Grub)	2.61×10 ^ 9				[51]
Dog (Yorkshire Terrier)	2.64×10 ^ 9				[51]
Springbok	two.72×10 ^ ix				[52]
Canis familiaris (Westward Highland White Terrier)	ii.83×ten ^ 9				[51]
Blesbok	3.06×10 ^ 9				[52]
Blue-and-yellow macaw	3.136×10 ^ ix		Brain only		[44]
Common squirrel monkey	3.246×x ^ ix				[47]
Crab-eating macaque	iii.44×ten ^ ix				[46]
Dog (High german Shepherd)	3.53×10 ^ 9				[51]
Tufted capuchin	3.691×10 ^ ix				[47]
Bonnet macaque	3.78×10 ^ nine				[46]
Striped hyena	3.885×x ^ 9				[48]
Dog (Komondor)	3.99×x ^ ix				[51]
Dog (Transylvanian Hound)	4.39×10 ^ 9				[51]
Panthera leo	iv.667×10 ^ 9				[48]
Greater kudu	4.91×10 ^ nine				[52]
Rhesus macaque	6.376×10 ^ 9				[47]
Chocolate-brown bear	ix.586×10 ^ 9				[48]
Giraffe	1.075×10 ^ 10				[52]
Xanthous baboon	ane.095×10 ^ 10				[46]
Chimpanzee	ii.eight×10 ^ x				[53]
Orangutan	3.26×10 ^ 10				[54]
Gorilla	iii.34×ten ^ x				[54]
Human	viii.half-dozen×10 ^ x	~1.5×x ^ 14	Neurons for average adult		[55] [56] [57]
African elephant	2.57×10 ^ 11				[58] [59]

List of animal species past forebrain (cerebrum or pallium) neuron number [edit]

The question of what concrete characteristic of an animate being makes an fauna intelligent has varied over the centuries. Ane early speculation was brain size (or weight, which provides the same ordering.) A second proposal was brain-to-body-mass ratio, and a tertiary was encephalization caliber, sometimes referred to as EQ. The electric current all-time predictor is number of neurons in the forebrain, based on Herculano-Houzel'southward improved neuron counts.^[i] It accounts most accurately for variations in dependence on the cerebellum. The elephant depends on its exceptionally large cerebellum, while birds make do with a much smaller one.

Differing methods have been used to count neurons and these may differ in degree of reliability. These are the optical fractionator, an awarding of stereology and the isotropic fractionator, a contempo methodological innovation. Most numbers in the list are the result of studies using the newer isotropic fractionator.^{[i] [37] [26] [41] [42] [44] [45] [46] [47] [52] [53] [54] [55] [56] [58] [sixty]} A variation of the optical fractionator was responsible for the previous total human brain neuron count of 100,000,000,000 neurons, which has been revised downwards to 86,000,000,000 by the utilise of the isotropic fractionator. This is in office why it may be considered to be less reliable. Finally, some numbers are the result of estimations based on correlations observed between number of cortical neurons and encephalon mass inside closely related taxa.

The following table gives information on the number of neurons estimated to exist in the sensory-associative structure (such as the pallium).

Common proper name	Boilerplate number of sensory-associative structure neurons[a]	Intraspecific variation[b]	Method[c]	Sensory-associative structure	Binomial classification	Epitome	Source
Mutual fruit wing	2,500*		Optical fractionator	Corpora pedunculata	Drosophila melanogaster		[61]
House cricket	50,000*		Optical fractionator	Corpora pedunculata	Acheta domesticus		[61]
Love bee	170,000*		Optical fractionator	Corpora pedunculata	Genus: Apis		[61]
Common cockroach	200,000*		Optical fractionator	Corpora pedunculata	Genus: Periplaneta		[61]
African sheath-tailed bat	5,000,000		Isotropic fractionator	Pallium (cortex)	Coleura afra		[38]
Rufous trident bat	half-dozen,000,000		Isotropic fractionator	Pallium (cortex)	Triaenops persicus		[38]
Naked mole-rat	vi,000,000	± 1,065,587[40]	Isotropic fractionator	Pallium (cortex)	Heterocephalus glaber		[37]
Petty free-tailed bat	6,000,000		Isotropic fractionator	Pallium (cortex)	Chaerephon pumilus		[38]
Common aptitude-fly bat	half dozen,000,000		Isotropic fractionator	Pallium (cortex)	Miniopterus schreibersii		[38]
Commerson's roundleaf bat	eight,000,000		Isotropic fractionator	Pallium (cortex)	Hipposideros commersoni		[38]
Heart-nosed bat	ten,000,000		Isotropic fractionator	Pallium (cortex)	Cardioderma cor		[38]
Ansell's mole-rat	10,000,000		Isotropic fractionator	Pallium (cortex)	Fukomys anselli		[43]
Smoky shrew	ten,000,000	± 352,000[40]	Isotropic fractionator	Pallium (cortex)	Sorex fumeus		[40]
Brusk-palated fruit bat	10,000,000		Isotropic fractionator	Pallium (cortex)	Casinycteris argynnis		[38]
Mashona mole-rat	12,000,000		Isotropic fractionator	Pallium (cortex)	Fukomys darlingi		[43]
Woermann's bat	12,000,000		Isotropic fractionator	Pallium (cortex)	Megaloglossus woermanni		[38]
Northern short-tailed shrew	12,000,000	± i,569,000[40]	Isotropic fractionator	Pallium (cortex)	Blarina brevicauda		[40]
Zenker's fruit bat	13,000,000		Isotropic fractionator	Pallium (cortex)	Scotonycteris zenkeri		[38]
House mouse	fourteen,000,000	± 2,242,257[twoscore]	Isotropic fractionator	Pallium (cortex)	Mus musculus		[40]
Hairy-tailed mole	16,000,000	± two,611,000[forty]	Isotropic fractionator	Pallium (cortex)	Parascalops breweri		[xl]
Star-nosed mole	17,000,000	± 3,105,000[forty]	Isotropic fractionator	Pallium (cortex)	Condylura cristata		[40]
Aureate hamster	17,000,000	± 3,619,934[40]	Isotropic fractionator	Pallium (cortex)	Mesocricetus auratus		[twoscore]
Diamond dove	18,209,000		Isotropic fractionator	Pallium (DVR)	Geopelia cuneata		[31]
King quail	twenty,523,000		Isotropic fractionator	Pallium (DVR)	Synoicus chinensis		[31]
Damaraland mole-rat	21,000,000		Isotropic fractionator	Pallium (cortex)	Fukomys damarensis		[43]
Hottentot golden mole	22,000,000	± 2,154,000[40]	Isotropic fractionator	Pallium (cortex)	Amblysomus hottentotus		[40]
Gray mouse lemur	22,310,000		Isotropic fractionator	Pallium (cortex)	Microcebus murinus		[46]
Common quail	22,568,000		Isotropic fractionator	Pallium (DVR)	Coturnix coturnix		[31]
Mechow'due south mole-rat	23,000,000		Isotropic fractionator	Pallium (cortex)	Fukomys mechowii		[43]
Franquet'due south epauletted fruit bat	23,000,000		Isotropic fractionator	Pallium (cortex)	Epomops franqueti		[38]
Hammer-headed bat	24,000,000		Isotropic fractionator	Pallium (cortex)	Hypsignathus mostrosus		[38]
Hedgehog	24,000,000^		Estimated	Pallium (cortex)	Subfamily Erinaceinae, various species		[62]
Silverish mole-rat	25,000,000		Isotropic fractionator	Pallium (cortex)	Heliophobius argenteocinereus		[43]
Wahlberg's epauletted fruit bat	26,000,000		Isotropic fractionator	Pallium (cortex)	Epomophorus wahlbergi		[38]
Cape mole-rat	26,000,000		Isotropic fractionator	Pallium (cortex)	Georychus capensis		[43]
Guyenne spiny rat	26,000,000	± 2,155,723[40]	Isotropic fractionator	Pallium (cortex)	Proechimys cayennensis		[twoscore]
Eastern rock elephant shrew	26,000,000	± 4,020,000[twoscore]	Isotropic fractionator	Pallium (cortex)	Elephantulus myurus		[40]
Eastern mole	27,000,000	± 5,113,000[40]	Isotropic fractionator	Pallium (cortex)	Scalopus aquaticus		[twoscore]
Opossum	27,000,000^		Estimated	Pallium (cortex)	Didelphis virginiana		[62]
Egyptian fruit bat	29,000,000		Isotropic fractionator	Pallium (cortex)	Rousettus aegyptiacus		[38]
Brown Rat	31,000,000	+ three,034,654[40]	Isotropic fractionator	Pallium (cortex)	Rattus norvegicus		[xl]
4-toed elephant shrew	34,000,000	± 5,840,000[forty]	Isotropic fractionator	Pallium (cortex)	Petrodromus tetradactylus		[forty]
Gray partridge	36,882,000		Isotropic fractionator	Pallium (DVR)	Perdix perdix		[31]
Ferret	38,950,000		Isotropic fractionator	Pallium (cortex)	Mustela putorius furo		[48]
Northern bobwhite	39,112,000		Isotropic fractionator	Pallium (DVR)	Colinus virginianus		[31]
African collared dove	39,997,000		Isotropic fractionator	Pallium (DVR)	Streptopelia roseogrisea		[31]
Cape dune mole-rat	43,000,000		Isotropic fractionator	Pallium (cortex)	Bathyergus suillus		[43]
Guinea pig	43,510,000	± 3,169,924[twoscore]	Isotropic fractionator	Pallium (cortex)	Cavia porcellus		[37]
Quebracho crested tinamou	48,292,000		Isotropic fractionator	Pallium (DVR)	Eudromia formosa		[31]
Mutual wood pigeon	51,325,000		Isotropic fractionator	Pallium (DVR)	Columba palumbus		[31]
Elegant crested tinamou	51,384,000		Isotropic fractionator	Pallium (DVR)	Eudromia elegans		[31]
Eurasian blackcap	52,000,000		Isotropic fractionator	Pallium (DVR)	Sylvia atricapilla		[44]
Prairie domestic dog	53,770,000	± 6,044,322[40]	Isotropic fractionator	Pallium (cortex)	Genus:Cynomys		[37]
Zebra finch	55,000,000		Isotropic fractionator	Pallium (DVR)	Taeniopygia guttata		[44]
Chilean tinamou	59,130,000		Isotropic fractionator	Pallium (DVR)	Nothoprocta perdicaria		[31]
Common treeshrew	lx,000,000	± 26,510,000[xl]	Isotropic fractionator	Pallium (cortex)	Tupaia glis		[40]
Red junglefowl	61,000,000		Isotropic fractionator	Pallium (DVR)	Gallus gallus		[44]
Goldcrest	64,000,000		Isotropic fractionator	Pallium (DVR)	Regulus regulus		[44]
European rabbit	71,450,000		Isotropic fractionator	Pallium (cortex)	Oryctolagus cuniculus		[37]
Rock pigeon	72,000,000		Isotropic fractionator	Pallium (DVR)	Columba livia		[44]
Eastern grayness squirrel	77,330,000	± two,634,444[40]	Isotropic fractionator	Pallium (cortex)	Sciurius carolinensis		[37]
Reeves's pheasant	80,688,000		Isotropic fractionator	Pallium (DVR)	Syrmaticus reevesii		[31]
Great tit	83,000,000		Isotropic fractionator	Pallium (DVR)	Parus major		[44]
Eurasian sparrowhawk	87,832,000		Isotropic fractionator	Pallium (DVR)	Accipiter nisus		[31]
Western tree hyrax	99,000,000		Isotropic fractionator	Pallium (cortex)	Dendrohyrax dorsalis		[40]
Green-rumped parrotlet	103,000,000		Isotropic fractionator	Pallium (DVR)	Forpus passerinus		[44]
Wild turkey	105,654,000		Isotropic fractionator	Pallium (DVR)	Meleagris gallopavo		[31]
Mallard	112,255,000		Isotropic fractionator	Pallium (DVR)	Anas platyrhynchos		[31]
Black-rumped agouti	113,000,000	± 2,576,768[40]	Isotropic fractionator	Pallium (cortex)	Dasyprocta prymnolopha		[i]
Banded mongoose	115,770,000		Isotropic fractionator	Pallium (cortex)	Mungos mungo		[48]
Victoria crowned pigeon	118,445,000		Isotropic fractionator	Pallium (DVR)	Goura victoria		[31]
Yellow-knobbed curassow	124,624,000		Isotropic fractionator	Pallium (DVR)	Crax daubentoni		[31]
Indian peafowl	129,621,000		Isotropic fractionator	Pallium (DVR)	Pavo cristatus		[31]
Marbled duck	130,142,000		Isotropic fractionator	Pallium (DVR)	Marmaronetta angustirostris		[31]
Pygmy falcon	131,898,000		Isotropic fractionator	Pallium (DVR)	Polihierax semitorquatus		[31]
Common blackbird	136,000,000		Isotropic fractionator	Pallium (DVR)	Turdus merula		[44]
Woods duck	138,206,000		Isotropic fractionator	Pallium (DVR)	Aix sponsa		[31]
Crimson-breasted goose	148,617,000		Isotropic fractionator	Pallium (DVR)	Branta ruficollis		[31]
Budgerigar	149,000,000		Isotropic fractionator	Pallium (DVR)	Melopsittacus undulatus		[44]
Eurasian teal	167,287,000		Isotropic fractionator	Pallium (DVR)	Anas crecca		[31]
Rock hyrax	198,000,000	± 29,082,000[xl]	Isotropic fractionator	Pallium (cortex)	Procavia capensis		[40]
Northern greater galago	226,000,000	± 87,570,000[40]	Isotropic fractionator	Pallium (cortex)	Otolemur garnettii		[40]
Common starling	226,000,000		Isotropic fractionator	Pallium (DVR)	Sturnus vulgaris		[44]
Common kestrel	237,903,000		Isotropic fractionator	Pallium (DVR)	Falco tinnunculus		[31]
Raccoon dog	240,180,000		Isotropic fractionator	Pallium (cortex)	Nyctereutes procyonoides		[51]
Common marmoset	245,000,000	± 81,180,000[40]	Isotropic fractionator	Pallium (cortex)	Callithrix jacchus		[forty]
House cat	249,830,000		Isotropic fractionator	Pallium (cortex)	Felis catus		[48]
Brown deport	250,970,000		Isotropic fractionator	Pallium (cortex)	Ursus arctos		[48]
Cockatiel	258,000,000		Isotropic fractionator	Pallium (DVR)	Nymphicus hollandicus		[44]
Greylag goose	258,650,000		Isotropic fractionator	Pallium (DVR)	Anser anser		[31]
Capybara	306,500,000	± 62,726,120[40]	Isotropic fractionator	Pallium (cortex)	Hydrochoerus hydrochaeris		[37]
Tarsius	310,000,000		Estimated	Pallium (cortex)	Genus: Tarsius, unknown species		[63]
Mute swan	323,661,000		Isotropic fractionator	Pallium (DVR)	Cygnus olor		[31]
Greater rhea	330,342,000		Isotropic fractionator	Pallium (DVR)	Rhea americana		[31]
Eastern rosella	333,000,000		Isotropic fractionator	Pallium (DVR)	Platycercus eximius		[44]
Common buzzard	351,700,000		Isotropic fractionator	Pallium (DVR)	Buteo buteo		[31]
Red fox	355,010,000		Isotropic fractionator	Pallium (cortex)	Vulpes vulpes		[51]
Goeldi's marmoset	357,130,000		Isotropic fractionator	Pallium (cortex)	Callimico goeldii		[46]
Eurasian pygmy owl	364,000,000		Isotropic fractionator	Pallium (DVR)	Glaucidium passerinum		[31]
Western greyness kangaroo	370,170,000		Isotropic fractionator	Pallium (cortex)	Macropus fuliginosus		[64]
Golden jackal	393,620,000		Isotropic fractionator	Pallium (cortex)	Canis aureus		[51]
Monk parakeet	396,000,000		Isotropic fractionator	Pallium (DVR)	Myiopsitta monachus		[44]
Springbok	396,900,000		Isotropic fractionator	Pallium (cortex)	Antidorcas marsupialis		[52]
Black vulture	398,899,000		Isotropic fractionator	Pallium (DVR)	Coragyps atratus		[31]
Azure-winged magpie	400,000,000		Isotropic fractionator	Pallium (DVR)	Cyanopica cyanus		[44]
Fiddling owl	400,822,000		Isotropic fractionator	Pallium (DVR)	Athene noctua		[31]
Common colina myna	410,000,000		Isotropic fractionator	Pallium (DVR)	Gracula religiosa		[44]
Domesticated pig	425,000,000*		Optical fractionator	Pallium (cortex)	Sus scrofa domesticus		[65]
Barn owl	437,000,000		Isotropic fractionator	Pallium (DVR)	Tyto alba		[44]
Emu	439,000,000		Isotropic fractionator	Pallium (DVR)	Dromaius novaehollandiae		[44]
W Highland White Terrier	440,160,000		Isotropic fractionator	Pallium (cortex)	Canis lupus familiaris		[51]
3-striped night monkey	442,000,000	± 111,310,000[xl]	Isotropic fractionator	Pallium (cortex)	Aotus trivirgatus		[ane]
Saker falcon	442,946,000		Isotropic fractionator	Pallium (DVR)	Falco cherrug		[31]
Eurasian magpie	443,000,000		Isotropic fractionator	Pallium (DVR)	Pica pica		[44]
Raccoon	453,000,000		Isotropic fractionator	Pallium (cortex)	Procyon lotor		[60]
Grub Chow	471,500,000		Isotropic fractionator	Pallium (cortex)	Canis lupus familiaris		[51]
Common ostrich	479,410,000		Isotropic fractionator	Pallium (DVR)	Struthio camelus		[31]
Western jackdaw	492,000,000		Isotropic fractionator	Pallium (DVR)	Coloeus monedula		[44]
Striped hyena	495,280,000		Isotropic fractionator	Pallium (cortex)	Hyaena hyaena		[48]
Chihuahua	513,330,000		Isotropic fractionator	Pallium (cortex)	Canis lupus familiaris		[51]
Eurasian jay	529,000,000		Isotropic fractionator	Pallium (DVR)	Garrulus glandarius		[44]
White-tailed eagle	542,926,000		Isotropic fractionator	Pallium (DVR)	Haliaeetus albicilla		[31]
Panthera leo	545,240,000		Isotropic fractionator	Pallium (cortex)	Panthera leo		[48]
Blesbok	570,670,000		Isotropic fractionator	Pallium (cortex)	Damaliscus pygargus phillipsi		[52]
Yorkshire Terrier	572,140,000		Isotropic fractionator	Pallium (cortex)	Canis lupus familiaris		[51]
Alexandrine parakeet	575,000,000		Isotropic fractionator	Pallium (DVR)	Psittacula eupatria		[44]
Tanimbar corella	599,000,000		Isotropic fractionator	Pallium (DVR)	Cacatua goffiniana		[44]
Golden retriever	627,000,000		Isotropic fractionator	Pallium (cortex)	Canis lupus familiaris		[48]
Gracile capuchin monkey	650,000,000^		Estimated	Pallium (cortex)	Genus: Cebus		[66]
Long-eared owl	673,000,000		Isotropic fractionator	Pallium (DVR)	Asio otus		[31]
Transylvanian Hound	725,760,000		Isotropic fractionator	Pallium (cortex)	Canis lupus familiaris		[51]
Greater kudu	762,570,000		Isotropic fractionator	Pallium (cortex)	Tragelaphus strepsiceros		[52]
Crab-eating macaque	800,960,000		Isotropic fractionator	Pallium (cortex)	Macaca fascicularis		[46]
Rook	820,000,000		Isotropic fractionator	Pallium (DVR)	Corvus frugilegus		[44]
Beagle	844,410,000		Isotropic fractionator	Pallium (cortex)	Canis lupus familiaris		[51]
Grey parrot	850,000,000		Isotropic fractionator	Pallium (DVR)	Psittacus erithacus		[44]
Komondor	883,380,000		Isotropic fractionator	Pallium (cortex)	Canis lupus familiaris		[51]
High german Shepherd	885,460,000		Isotropic fractionator	Pallium (cortex)	Canis lupus familiaris		[51]
Sulphur-crested cockatoo	1,135,000,000		Isotropic fractionator	Pallium (DVR)	Cacatua galerita		[44]
Tufted capuchin	1,140,000,000		Isotropic fractionator	Pallium (cortex)	Sapajus apella		[twoscore] [1]
Tawny owl	1,153,000,000		Isotropic fractionator	Pallium (DVR)	Strix aluco		[31]
Harp seal	1,168,000,000^ half dozen,100,000,000*		Estimated Optical fractionator	Pallium (cortex)	Pagophilus groenlandicus		[67] [68]
Horse	1,200,000,000*		Optical fractionator	Pallium (cortex)	Equus ferus caballus		[69]
Raven	one,204,000,000		Isotropic fractionator	Pallium (DVR)	Corvus corax		[44]
Snowy owl	1,270,000,000		Isotropic fractionator	Pallium (DVR)	Bubo scandiacus		[31]
Kea	1,281,000,000		Isotropic fractionator	Pallium (DVR)	Nestor notabilis		[44]
Eurasian hawkeye-owl	1,328,000,000		Isotropic fractionator	Pallium (DVR)	Bubo bubo		[31]
Common squirrel monkey	1,340,000,000	± xx,000,000[twoscore]	Isotropic fractionator	Pallium (cortex)	Saimiri sciureus		[40]
Bonnet macaque	1,660,000,000		Isotropic fractionator	Pallium (cortex)	Macaca radiata		[xl]
Rhesus macaque	one,710,000,000		Isotropic fractionator	Pallium (cortex)	Macaca mulatta		[twoscore]
Giraffe	one,731,000,000		Isotropic fractionator	Pallium (cortex)	Giraffa camelopardalis		[52]
Blueish and yellow macaw	1,900,000,000		Isotropic fractionator	Pallium (DVR)	Ara ararauna		[44]
Pygmy sperm whale	2,020,000,000*		Optical fractionator	Pallium (cortex)	Kogia breviceps		[70]
Leopard seal	ii,386,000,000^		Estimated	Pallium (cortex)	Hydrurga leptonyx		[67]
Guenon	2,500,000,000^		Estimated	Pallium (cortex)	Genus: Cercopithecus, unknown species		[63]
Pigtail Macaque	ii,531,000,000^		Estimated	Pallium (cortex)	Macaca nemestrina		[67]
Gelada birdie	2,568,000,000^		Estimated	Pallium (cortex)	Theropithecus gelada		[67]
Cerise-and-green macaw	2,646,000,000^		Estimated	Pallium (DVR)	Ara chloropterus		[67]
Harbor porpoise	2,750,000,000*		Optical fractionator	Pallium (cortex)	Phocoena phocoena		[67] [68]
Yellow baboon	2,880,000,000		Isotropic fractionator	Pallium (cortex)	Papio cynocephalus		[46]
Hyacinth macaw	2,944,000,000^		Estimated	Pallium (DVR)	Anodorhynchus hyacinthinus		[67]
Hamadryas baboon	two,990,000,000^		Estimated	Pallium (cortex)	Papio hamadryas		[67]
Mandrill	iii,102,000,000^		Estimated	Pallium (cortex)	Mandrillus sphinx		[67]
Walrus	3,929,000,000^		Estimated	Pallium (cortex)	Odobenus rosmarus		[67]
Southern elephant seal	three,994,000,000^		Estimated	Pallium (cortex)	Mirounga leonina		[67]
African elephant	5,600,000,000		Isotropic fractionator	Pallium (cortex)	Loxodonta africana		[58]
Curt-beaked mutual dolphin	6,700,000,000*		Optical fractionator	Pallium (cortex)	Delphinus delphis		[67] [70]
Asian elephant	6,775,000,000^		Estimated	Pallium (cortex)	Elephas maximus		[67]
Pygmy chimpanzee or bonobo	7,250,000,000^		Estimated	Pallium (cortex)	Pan paniscus		[67]
Chimpanzee	vii,400,000,000*		Optical fractionator	Pallium (cortex)	Pan troglodytes		[71]
Orangutan	7,704,000,000^ - 8,900,000,000^		Estimated	Pallium (cortex)	Genus: Pongo		[67] [54]
Western gorilla	9,100,000,000^		Estimated	Pallium (cortex)	Gorilla gorilla		[67] [54]
Cuvier'south beaked whale	nine,100,000,000*		Optical fractionator	Pallium (cortex)	Ziphius cavirostris		[70]
Beluga whale	10,000,000,000^		Estimated	Pallium (cortex)	Delphinapterus leucas		[67]
Bottlenose dolphin	12,700,000,000*		Optical fractionator	Pallium (cortex)	Tursiops truncatus		[67] [lxx]
Mutual minke whale	12,800,000,000*		Optical fractionator	Pallium (cortex)	Balaenoptera acutorostrata		[72]
Fin whale	fifteen,000,000,000*		Optical fractionator	Pallium (cortex)	Balaenoptera physalus		[67] [73]
Blueish whale	fifteen,000,000,000^		Estimated	Pallium (cortex)	Balaenoptera musculus		[67]
Man	sixteen,340,000,000 21,000,000,000*	± 2,170,000,000[twoscore]	Isotropic fractionator Optical fractionator	Pallium (cortex)	Human being sapiens		[40] [55] [1] [74] [75]
Risso's dolphin	18,750,000,000^		Estimated	Pallium (cortex)	Grampus griseus		[67]
Brusk-finned pilot whale	35,000,000,000^		Estimated	Pallium (cortex)	Globicephala macrorhynchus		[67]
Long-finned pilot whale	37,200,000,000*		Optical fractionator	Pallium (cortex)	Globicephala melas		[76]
Killer whale	43,100,000,000*		Optical fractionator	Pallium (cortex)	Orcinus orca		[67] [70]

See also [edit]

• Lists of animals
• Theory of mind in animals
• Brain size
• Brain-to-body mass ratio
• Encephalization quotient
• Connectome
• Connectomics
• Cranial chapters
• Noogenesis

Notes [edit]

1. ^ ^ = Estimated
   * = Optical fractionator
2. ^ ± standard difference
3. ^ For the estimated values, the numbers of cortical neurons estimated from brain mass for different mammalian and bird orders are based on correlation observed between number of cortical neuron and brain mass per order^[40]

References [edit]

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Source: https://en.wikipedia.org/wiki/List_of_animals_by_number_of_neurons

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